@article {layton2009drosophila, title = {Drosophila morphogenesis: tissue force laws and the modeling of dorsal closure}, journal = {HFSP journal}, volume = {3}, number = {6}, year = {2009}, pages = {441{\textendash}460}, publisher = {Taylor \& Francis}, author = {Layton, Anita T and Toyama, Yusuke and Yang, Guo-Qiang and Edwards, Glenn S and Kiehart, Daniel P and Venakides, Stephanos} }